Cyrtomium anomophyllum (Zenker) Standish ex J.J.Blandy, Proc. Roy. Hort. Soc. London 2: 364 (1862) (Syn: Aspidium anomophyllum Zenker; Aspidium caryotideum f. macropterum KunzeCyrtomium beddomei S.R.GhoshCyrtomium brevicuneatum Ching & Shing ; Cyrtomium caryotideum var. anomophyllum (Zenker) T.Moore; Cyrtomium kansuense Ching & ShingCyrtomium kungshanense Ching & ShingCyrtomium macrophyllum var. microindusium (Sa.Kurata) K.Iwats.Cyrtomium maximum Ching & ShingCyrtomium microindusium Sa.KurataCyrtomium neocaryotideum Ching & ShingCyrtomium nervosum Ching & ShingCyrtomium shunningense Ching & ShingCyrtomium yuanum Ching & ShingPhanerophlebia macroptera (Kunze) Akasawa ; Polystichum anomophyllum (Zenker) Nakai; Polystichum caryotideum var. macropterum (Kunze) NakaiPolystichum nervosum (Ching & Shing) C.M.Kuo);
Indian Subcontinent to China (W. Sichuan, NW. Yunnan), Japan (Kyushu), Taiwan: Assam, China South-Central, East Himalaya, India, Japan, Myanmar, Nepal, Pakistan, Taiwan, Tibet, West Himalaya as per POWO;

Fern for id Kalatope – Al040512:
A fern for id please…
Location Kalatope, Chamba
Altitude 2100 mts

Habitat wild
Season March

The Picture is of Cyrtomium caryotideum (Wall. Ex Hook & Grev) Presl

Cyrtomium anomophyllum, I think! It’s one of those specimens one occasionally gets that has developed a bit of an acute auricle – towards C. caryotideum. Then they can be rather difficult to say which of the two they are (but can’t be hybrids as both are triploid apomict so can’t hybridise). I notice also the little dark centres to the indusium, which are typical of anomophyllum. Syn.: C. nervosum, C. microindusium.
Nice photos showing the teeth and its matt surface texture – in strong contrast to the Japanese and E. Chinese species, C. falcatum, occasionally cultivated in India.

what had impressed me was the large “leaf” and enormous numbers of Sori many of which seem to have burst open and released the spores.

Incidentally, as there were two answers, what makes me think it is more likely C. anomophyllum than caryotideum is the lower leaf in the photo, showing the very rounded pinna-bases (as also the lowest pinna on the upper frond), and I have grown plants of it and seen how they can occasionally form a smallish, acute auricle in mid and upper pinnae (approaching the narrow auricles of caryotideum, but usually less prominent). Also the dark centres to the indusia. But it’s true to say those two are a bit difficult sometimes, and alternatively sometimes C. anomophyllum can appear close to C. macrophyllum, but with teeth at least at the pinna-apices. They have long been recognised by the critical and carefulBotanists in Japan. Dr. Sadamu Matsumoto at Tsukuba has shown that all these species are hybrid-derived and has been able to identify some of the ancestral diploids – so some degree of occasional intermediacy in morphology is not so surprising really.

Well, I have included an updated Himachal and J. and K. list on the Indian Ferns website ( You open the Home Page, go to New Members and join it (free), then you can open up Files on the left, and my paper on Nepal ferns etc. gives Himalayan State lists. This is soon to be updated to include other States, until all Indian States’ pteridophytes are listed. But I am a little busy at present and can’t do the expansion for a couple of months at least, though it is half ready.
… at the Himalayan Botanical Research Institute, Palampur, is also working on Himachal ferns, as has … at Panjab University, Chandigarh, for many decades.

Cyrtomium sp. ABJUL01/33 : 6 posts by 3 authors. 1 image.
I found this in a dark understorey on a slope. I could not reach close enough to the fern for a closer look at the scales but could stretch my arm to touch it. It had the feel of a thick dry leaf and had no spores underneath. I am not sure if we can tell the species from looking at the frond only but it could be Cyrtomium nervosum. Please advise.
Above Mcleodganj, Dharamshala, HP
19 July 2016

Yes, That’s Cyrtomium anomophyllum (syn.: C. nervosum) – occasionally it gets that attractive dark stripe along the pinna midribs.

Found the sori on the Cyrtomium anomophyllum and am sharing them with you.
Above Dal Lake, Dharamshala, HP

13-15 March 2017
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