Swertia nervosa

Swertia nervosa (Wall. ex G. Don) C. B. Cl., Fl. Brit. India 4(10): 125 1883. (syn: Agathotes nervosa Wall. ex G. Don; Ophelia nervosa (Wall. ex G. Don) Griseb.; Swertia cavaleriei H. Lév.; Swertia nervosa Wall. (ambiguous synonym));
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Bhutan, Darjeeling, Sikkim, India, Nepal, China (SE-Gansu, Guangxi, Guizhou, SW-Shaanxi, Sichuan, Yunnan), Tibet as per Catalogue of Life;

Keys as per eFI thread:
Flowers are nearly similar in Swertia alata and Swertia nervosa. Basic difference is leaf sessile and base cordate in S. alata and leaf petiolate and base cuneate in S. nervosa, apart from other micro morphological characteristics.
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Swertia nervosa (Wall. ex G. Don) C. B. Cl. (accepted name) ?? : 4 posts by 2 authors. Attachments (10)

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Thanks, … Appears close as per images at

This is Swertia nervosa as per keys and discussions at Swertia nervosa (Wall. ex G.Don) C.B.Clarke

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Swertia nervosa (Wall. ex G.Don) C.B.Clarke:
Location: Godam, Chandragiri, Kathmandu, Nepal
Altitude : 1612m.
Date: 31 October 2023
Habit : Wild

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Nepali Names : तिते चिराइतो Tite Chiraaito / कालो चिराइतो Kaalo Chiraaito

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How is it different from
https://efloraofindia.com/2020/10/09/swertia-alata/?

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I guess there is a slight difference in the floral pattern/ design !

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Can you find out the keys as both the species are distributed in your area and Bhutan and NE India as per POWO:
https://powo.science.kew.org/names:370998-1
https://powo.science.kew.org/names:370786-1

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This is Swertia nervosa only as per my knowledge.

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May I request you to provide the differences between these two species (and keys).

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Sir flowers are nearly similar in Swertia alata and Swertia nervosa. Basic difference is leaf sessile and base cordate in S. alata and leaf petiolate and base cuneate in S. nervosa, apart from other micro morphological characteristics.

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Do you consider the posted lower leaves as petiolate with cuneate base?

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References:
Catalogue of Life  The Plant List Ver.1.1  Tropicos  Flora of China  FOC illustration  Annotated checklist of Flowering plants of Nepal  EOL (with specimen)  India Biodiversity Portal  Swertia in Nepal – Gentian Research Network – Rutgers University  ResearchGate