Veronica biloba Schreb. ex L., Mant. Pl. 2: 172 172 1771. (Syn: Pocilla biloba (L.) W.A.Weber; Veronica biloba var. densa G.H.Dar & Kachroo; Veronica chantavica Pavlov; Veronica exilis H.Schott ex Boiss.; Veronica microtheca Boiss. & Balansa; Veronica nevskii Boriss.; Veronica perpusilla Nevski);
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Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, SSW-Anatolia, SW-Anatolia, W-Anatolia), Iran (EC-Iran, E-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran), Iraq (NE-Iraq), Lebanon (Antilebanon, C-Lebanon), Sinai peninsula (S-Sinai), China (Gansu, Nei Mongol, Ningxia, Qinghai, Shaanxi, Sichuan, Xinjiang), Tibet, Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan, Faryab, Ghazni, Herat, Kabul, Kunar / Nuristan, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan, Samangan, Takhar), India, Kazakhstan, Kyrgyzstan, Mongolia, Nepal, Pakistan (Baluchistan, Kurram, Chitral, Swat, Hazara, Murree), Pakistani Kashmir (Gilgit, Baltistan, Deosai), Jammu & Kashmir (Ladakh, Poonch, Kashmir), Tajikistan, Turkmenistan, Uzbekistan, Northern Caucasus, Armenia, Azerbaijan, Georgia [Caucasus], Siberia (W-Siberia), Russian Far East (I), USA (I) (Colorado (I), Idaho (I), Illinois (I), Kansas (I), Montana (I), Nebraska (I), Nevada (I), New York (I), Oregon (I), Utah (I), Washington State (I), Wyoming (I)), Canada (I) (British Columbia (I)) as per Catalogue of Life;
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‘Twolobe Speedwell’;
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Scrophulariaceae of the Western Himalayas By Francis . Pennell (Description & Keys- Veronica capitata, cachemirica, lanosa, macrostemon, lasiocarpa, serpyllifolia, verna, arvensis, didyma (syn. of Veronica polita Fr. as per The Plant List), persica, perpusilla (syn. of Veronica pusilla Kotschy & Boiss. as per The Plant List), uncinata (syn. of Veronica rubrifolia Boiss. as per The Plant List), biloba, campylopoda, stewartii, koelzii, cephaloides (syn. of Veronica ciliata subsp. cephaloides (Pennell) D.Y. Hong as per The Plant List), hirta, nana (syn. of Veronica ciliata subsp. cephaloides (Pennell) D.Y. Hong as per The Plant List), melissaefolia, javanica, cana, umbelliformis (syn. of Veronica szechuanica subsp. sikkimensis (Hook. f.) D.Y. Hong as per The Plnat List), beccabunga, anagallis-aquatica, secunda, salina & undulata)
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Erect annual herb usually branched from base; leaves oblong to ovate-lanceolate, up to 3 cm long, margin shallowly dentate; racemes mostly terminal, sparsely glandular hairy; bracts smaller than leaves, entire; calyx 4-lobed, lobes ovate-lanceolate, 4-8 mm long in fruit with entire margin, 3-veined; corolla white or blue-purple, 3-4 mm diam.; capsule strongly compressed, shorter than calyx, deeply notched, lobes appearing almost free; style very short, included in notch. 
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Scrophulariaceae Fortnight: Veronica for id from Chakrata-NS 12 : Attachments (6). 4 posts by 3 authors.
This Veronica species was shot from Budher caves in Chakrata, please help to id.. can this be Veronica biloba?
Now placed in Plantaginaceae


Id already confirmed as per Flora of Chakrata 2012: For id from Budher caves:


Yes Veronica biloba. Nice photographs


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Flora of Chakrata 2012: For id from Budher caves:
This very small herb was shot from meadows of Budher caves on May 17, 2012. I hope this is a Veronica species…
wish to have exact id…
thanks in advance..

Veronica biloba I hope


I googled Veronica biloba and all images seem to have serrate leaves. Is the leaf variation normal?


Please go through these.
http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200021307
http://www.efloras.org/object_page.aspx?object_id=3821&flora_id=2



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Scrophulariaceae Fortnight: Veronica biloba from Kashmir -GS-48 :  (Mixed thread): 2 correct images.. 2 posts by 2 authors.
Veronica biloba Linnaeus, Mant. Pl. 2: 172. 1771.
Erect annual herb usually branched from base; leaves oblong to ovate-lanceolate, up to 3 cm long, margin shallowly dentate; racemes mostly terminal, sparsely glandular hairy; bracts smaller than leaves, entire; calyx 4-lobed, lobes ovate-lanceolate, 4-8 mm long in fruit with entire margin, 3-veined; corolla white or blue-purple, 3-4 mm diam.; capsule strongly compressed, shorter than calyx, deeply notched, lobes appearing almost free; style very short, included in notch.
Common as weed in fields and roadsides. Photographed from Gulmarg, Kashmir.

Thanks sir for this upload, I have shared this from Chakrata area.. perhaps flowers in April-May, could not find this during present tour to the same place..

A feedback from eFI page:
“In the end picture of set 2 above, you’ll see all the capsules are nodding (bending down). You’ll see the others pictures do not have this, and Flora of Turkey states explicitly that V. biloba capsules do not nod, and the illustration you provide likewise does not show nodding (http://www.efloras.org/object_page.aspx?object_id=3821&flora_id=2) . So I would suggest that picture is not V. biloba.
(There are certainly Veronica in Turkey with deeply split capsules which do nod, such as Veronica campylopoda, presumably there will be others taken across a wider geography.)”


Yes …, now I see my collections from Kashmir are a mix of two

First and third image with recurved pedicels, nodding capsules, capsules, covered with glandular hairs, longer style is C. campylopoda
2nd and 4th image with spreading pedicels,  erect capsules covered with short hairs and shorter style is V. biloba
They are from different locations. I will upload them separately with clearer high resolution images with locations.

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Veronica biloba Linnaeus, Mant. Pl. 2: 172. 1771.
Erect annual herb usually branched from base; leaves oblong to ovate-lanceolate, up to 3 cm long, margin shallowly dentate; racemes mostly terminal, sparsely glandular hairy; bracts smaller than leaves, entire; calyx 4-lobed, lobes ovate-lanceolate, 4-8 mm long in fruit with entire margin, 3-veined; corolla white or blue-purple, 3-4 mm diam.; capsule strongly compressed, shorter than calyx, deeply notched, lobes appearing almost free; style very short, included in notch.
Common as weed in fields and roadsides.
Photographed
from Gulmarg.

This herb is present in my database in section of Healing Flora of Ladakh. Veronica biloba based Formulations are used for Insomnia.



Veronica biloba from Gulmarg Kashmir-30042023-3: 4 high res. images.
Veronica biloba Linn.

The species is very similar and often confused with V. campylopoda, but can be differentiated as under:
1. Calyx lobes narrower (nearly 1.6 mm broad in fruit), pedicel in fruit distinctly recurved with nodding capsules, capsule covered with long hairs, some gland-tipped,lobes separated almost to the base (united for less than 0.5 mm), style 1-1.5 mm long, seeds transversally rugose, 8-14 per capsule …….V. campylopoda
1. Calyx lobes broader (2-4 mm broad in fruit), pedicel in fruit spreading, capsule not nodding, puberulent usually without gland-tipped hairs, separated 1/2 to 2/3 (united 0.5-1 mm at base), style 0,5-0,8 mm long, seeds obscurely rugose, less than 8 per capsule …….V. biloba
V. biloba photographed from Gulmarg, Kashmir, Alt. 2500 m.




Veronica biloba from Pahalgam Kashmir-30042023-4: 3 high res. images.
Veronica biloba Linn.

The species is very similar and often confused with V. campylopoda, but can be differentiated as under:
1. Calyx lobes narrower (nearly 1.6 mm broad in fruit), pedicel in fruit distinctly recurved with nodding capsules, capsule covered with long hairs, some gland-tipped,lobes separated almost to the base (united for less than 0.5 mm), style 1-1.5 mm long, seeds transversally rugose, 8-14 per capsule …….V. campylopoda
1. Calyx lobes broader (2-4 mm broad in fruit), pedicel in fruit spreading, capsule not nodding, puberulent usually without gland-tipped hairs, separated 1/2 to 2/3 (united 0.5-1 mm at base), style 0,5-0,8 mm long, seeds obscurely rugose, less than 8 per capsule …….V. biloba 

Photographed from Pahalgam, Kashmir, alt. 2600 m



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References:

Comments

  1. David M

    In the end picture of set 2 above, you’ll see all the capsules are nodding (bending down). You’ll see the others pictures do not have this, and Flora of Turkey states explicitly that V. biloba capsules do not nod, and the illustration you provide likewise does not show nodding ( http://www.efloras.org/object_page.aspx?object_id=3821&flora_id=2 ) . So I would suggest that picture is not V. biloba.
    (There are certainly Veronica in Turkey with deeply split capsules which do nod, such as Veronica campylopoda, presumably there will be others taken across a wider geography.)
    David

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