Duhaldea eupatorioides

Duhaldea eupatorioides (Wall. ex DC.) A. A. Anderberg, Pl. Syst. Evol. 176: 104 (1991) (syn: Amphirhapis heterotricha DC.; Aster zayuensis Y.L.Chen; Conyza eupatorioides Wall.; Conyza heterotricha Wall.; Helenium eupatoriodes (DC.) Kuntze; Inula eupatorioides DC.; Solidago heterotricha Wall.);
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Himalaya to SE. Tibet and Indo-China: Assam, East Himalaya, Laos, Myanmar, Nepal, Pakistan, Tibet, Vietnam, West Himalaya as per POWO;
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India (Uttar Pradesh, Sikkim, Nagaland, Meghalaya, Jammu), Nepal, Bhutan, Pakistan (Dir, Salt Range), SE-Tibet, Laos, Myanmar [Burma], Vietnam as per CoL;
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Common name: Boneset-Leaf Inula
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Woolly shrubs, up to 3 m tall, stout; branches pubescent. Leaves simple, alternate, spiral; lamina ca. 3.5-25 x 1.2-7.5 cm, lanceolate or elliptic-oblong, acute or rounded at base, acuminate at apex, margins irregularly dentate, scaberulous above, pubescent beneath. Heads up to ca. 0.8 cm across, numerous in terminal or upper axillary corymbs; involucral bracts multiseriate, subulate, tomentose; florets bright yellow. Achenes narrowly oblong, adpressed silky-pubescent; pappus straw yellow or reddish with brown tinge, plumose.
Flowering & Fruiting: October-February.
Grassy areas
(Attributions- French Institute of Pondicherry as per India Biodiversity Portal)
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MS Dec,2018/04 Asteraceae for ID : 7 posts by 2 authors. Attachments (2)
Location : Tan, Mizoram

Date : 06-12-2018
Habit : Small shrub
Habitat : Wild

Does not appear to be a Senecioneae member.

To me appears to be some Duhaldea‎ species, but not matching with the existing species herein as per comparative images.
Pl. check Duhaldea eupatorioides as per research gate link one and two
Senecio. But currently do not have the flora to reach spp.
The plant in pdf is not Senecio its Duhaldea cuspidata
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MS Dec,2018/09 Asteraceae (Duhaldea sp.?) for Id. : 7 posts by 3 authors. Attachments (3)
Location : Tan, Mizoram
Altitude : 1600 – 1,900 m.
Date : 06-12-2018
Habit : Shrub
Habitat : Wild
Does not match with any species available so far as per comparative images at

Also does not match as per comparative images at
/species/a—l/ar/asteraceae/asteroideae/inuleae

Senecio sp. ??

Duhaldea cuspidata as per another thread: MS Dec,2018/04 Asteraceae for ID

This appears more closer to images and details at Synotis cappa (Buch.-Ham. ex D.Don) C.Jeffrey & Y.L.Chen

Duhaldea cuspidata.

Pl. see at Synotis cappa (Buch.-Ham. ex D.Don) C.Jeffrey & Y.L.Chen

I think it is not Duhaldea cuspidata because of Duhaldea cuspidata has stalked leaves, and the leaves in this plant are virtually stalkless. Synotis cappa appears close but if you look carefully, in Synotis cappa the ray florets are longer than the central disk of the flower-heads. In this plant the ray florets are much shorter than the central disk.
I think it is Duhaldea eupatorioides. The structure of the inflorescence and the leaves agree with the illustration attached here, and to the description in this paper.

I do not think so.
Pl. see
https://efloraofindia.com/2021/06/17/duhaldea-eupatorioides/
Pl. see detailed description as given below:

Description from Flora of China:
Synotis cappa (Buchanan-Hamilton ex D. Don) C. Jeffrey & Y. L. Chen		 Duhaldea eupatorioides (Wall. ex DC.) Anderb. as per The genus Duhaldea (Asteraceae) in India (2020)
Subshrubs or shrubby herbs, rhizomatous. Stems erect, to 150 cm tall, often flexuous, densely lanate or arachnoid-tomentose, ± glabrescent when old, simple or usually branching above in synflorescences, leafless in lower part at anthesis.

Leaves petiolate or sometimes subsessile; petiole to 10 mm, stout, densely tomentose, base often auriculate; blade broadly to narrowly obovate-oblanceolate or oblong-elliptic, 10-28 × 4-8 cm, papery, abaxially densely to sparsely fulvous pubescent on veins and densely to sparsely whitish tomentose, sometimes ± glabrescent, adaxially shortly densely to sparsely pubescent and sometimes thinly white arachnoid to subglabrous, pinnately veined, lateral veins 12-28, arcuate-ascending, base attenuate, margin finely to coarsely mucronulate-serrate, apically acuminate. Upper leaves and leaves of branches lanceolate or linear-lanceolate, gradually smaller.

Capitula radiate, usually numerous, arranged in terminal and axillary usually 10-20-headed dense compound corymbs or thyrses; peduncles 5-10 mm, densely tomentose, with a few linear or linear-subulate bracts.

Involucres narrowly campanulate, 5-7 × 2-4 mm, calyculate; bracts of calyculus ca. 8, linear-lanceolate, 3-5 mm, apically acuminate; phyllaries 8-13, linear-lanceolate, ca. 1 mm wide, herbaceous, densely tomentose, margin narrowly scarious, apically obtuse to acute and puberulent.

Ray florets ca. 8; corolla tube ca. 2.5 mm; lamina yellow, ca. 3.5 × 1 mm, 4-veined, 3-denticulate. Disk florets 11-17; corolla yellow, ca. 5 mm, with ca. 1.5 mm tube and funnelform limb; lobes ovate-oblong, ca. 1.5 mm, apically acute. Anthers ca. 2.5 mm; anther tails slightly exceeding antheropodia; appendages ovate-oblong; antheropodia rather slender, basally slightly dilated. Style branches ca. 1 mm, fringed with short papillae, terminal tuft not evident. Achenes ca. 2 mm, glabrous. Pappus white, ca. 5 mm.

Fl. Oct-Jan.

Forest margins, thickets, streamsides, meadows; 1500-2300 m.

Guangxi, Sichuan, Xizang, Yunnan [Bhutan, NE India, Myanmar, Nepal, N Thailand].

  

 

 Stout shrubs, 1–3 m tall. Stem pubescent, terete; branches brownish-purple.

Leaves spirally arranged, rigid, leathery,shortly petioled; lamina elliptic-oblong or lanceolate-acuminate, 5–25 × 1.5–7.5 cm, coriaceous, irregularly toothed, cuneate or rounded at base, dark green above, scabrid, pale green below, pubescent, dense towards nerves, sometimes glandular, lateral nerves prominent, shorter, with much-reticulated nervation beneath.Corymbs polycephalous. Heads terminal or in upper axillary panicles; capitula crowned with short peduncles, 0.5–1.0 cm in diam.

Involucral bracts 2–4 × 0.5–1.0 mm long, 4-seriate, lax, imbricate, lanceolate to linear-lanceolate, brownish, villous to velutinous, sometimes glandular at tip.


Ray florets a few, shortly radiate. Disc florets tubular, 5- toothed. Cypselae narrowly obovate–oblong, 1.5– 2.0 × 0.25–0.5 mm, yellowish brown, sparsely white pubescent, 8–10-ribbed. Pappus uniseriate, bristles 22–24, 4–5 mm long, cream-golden.

Flowering and fruiting from August to February.

Habitat: Grows on rocky slopes, between 1700– 1800 m elevations.

Distribution: Bhutan, China, India, Laos, Myanmar, Nepal, Pakistan, Thailand and Vietnam.
Thank you …i for the detailed comparison.
But I do not think the images at the eFI link correctly represent the species Duhaldea eupatorioides. The flowers seem to have almost no ray florets. I am attaching an image of Duhaldea eupatorioides from this paper. Another paper summarizes Duhaldea eupatorioides as follows:
 Notes: This species is similar to D. cuspidata, but differs in much stouter habit, shorter and stouter petioles, more rigid leaves with much reticulated nervation beneath and narrower ligules.
Comparing with the images of Duhaldea cuspida, I feel … images fit the description of Duhaldea eupatorioides reasonably well.

But description in the publication says Ray florets a few, shortly radiate for Duhaldea eupatorioides. Also see the image of it in the publication, which is closer to our images in eFI site.

This is getting more confusing. The leaves of Duhaldea eupatorioides should be remotely toothed, as is seen in herbarium image at POWO. The leaves depicted in the drawing in that paper are correctly represented (attached here). However the leaves in the photograph of the plant, in the same paper, look very different. The leaf-margins of … plant do agree with the herbarium images of POWO and the drawing attached here. The flowers of … plant do look similar to the flowers in Chen’s clear image of Duhaldea eupatorioides that I posted earlier.  For me that is a reasonable reason for considering it as Duhaldea eupatorioides.

https://ppbc.iplant.cn/tu/10919517
https://ppbc.iplant.cn/tu/10919511

These images on the iPlant link agree with my understanding of this species.

May I request you for post high res. images to see the phyllaries.


2 very high res. images.

Yes, I agree with the suggested id by …
Phyllaries seem to match as per these high res. image with those posted by …

 

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References:
POWO  Catalogue of Life  The Plant List Ver.1.1  GCC  GBIF (High resolution specimens) Annotated checklist of Flowering plants of Nepal  Flowers of India  Pan India Bouquets  India Biodiversity Portal
The genus Duhaldea (Asteraceae) in India – Shweta Shekhar, Arne Alfred Anderberg & Arun Pandey June 2020 Rheedea 30(2):257-269 (Abstract- Based on the field and herbarium data, six species of the genus Duhaldea (sensu stricto) are recognized in India. A key to the Indian species, brief descriptions and illustrations are provided along with data on phenology, distribution, ecology, chromosome numbers and ethnobotanical uses)