Keys in Flora of Bhutan:
1. Leaves simple (but often lobed) ……… .. 2
+ Leaves compound …………………………. .. 5
2. Tendrils usually twice bifid, never borne on peduncles; flowers 4-merous; style elongate ……….. .. 5. Cissus
+ Tendrils once-bifid, or with one branch again bifid, often borne on peduncles; flowers 5-merous; style very short or absent …….. .. 3
3. Flowers in rounded cymes: petals free and spreading at anthesis ……….  2. Ampelocissus
+ Flowers in panicles; petals free and spreading or united and shed as a cap at anthesis ……… .. 4
4. Stems, petioles and peduncles never softly glandular-bristly; petals united at apex and shed as a cap at anthesis …… 1. Vitis
+ Stems, petioles and peduncles softly glandular-bristly; petals free at apex and spreading at anthesis ……………………. .. 2. Ampelocissus (A. barbata)
5. Leaves trifoliolate ………………………………………… .. 6
+ Leaves palmately or pedately 4 — 7-foliolate …… .. 9
6. Young stems whitish woolly; lateral leaflets often lobed; peduncles usually bearing a tendril; flowers 5-merous …….. .. 2. Ampelocissus (A. divaricatus)
+ Stems glabrous or pubescent but never whitish woolly; lateral leaflets unlobed; peduncles without tendril; flowers 4- or 5-merous ……………. 7
7. Tendrils 2 – 3 x bifid, branches ending in disc-like pads; flowers in terminal and leaf-opposed cymes, 5-merous; style stout, shonly columnar  …………. 3. Parthenocissus
+ Tendrils simple or 1 — 2 x bifid, but branches not ending in disc-like pads; flowers in axillary or pseudo-axillary cymes, 4-merous; style absent or elongate and slender ……………….. .. 8
8. Tendrils simple; leaves glabrous beneath; stigma conspicuously 4-lobed or disc-like, sessile on ovary ………… .. 4. Tetrastigma
+ Tendrils 1 — 2 x bifid; leaves softly villous beneath; stigma minute on slender elongate style …………… .. 7. Cayratia (C. geniculata)
9. Leaves palmately compound; berry 1 – 2-seeded ……………. 10
+ Leaves pedately compound; berry 2 — 4-seeded …………. .. 12
10. Tendrils simple or radiately branched; style absent or very shon ……. 4. Tetrastigma
+ Tendrils 1 — 2 x bifid; style elongate ………………………………………….. .. 11
1 1. Young stems, petioles, inflorescences and leaves glabrous or almost so; leaflet margins sharply serrulate; cymes leaf-opposed; berry 2 — 2.5cm; seed c 16 x 10mm …………… .. 5.Cissus (C. elongata)
+ Young stems, petioles, inflorescences and leaves (beneath) softly pubescent; leaflet margins crenate; cymes pseudo-axillary or temtinal; berry c l.5cm diameter; seed c 8 x 6mm …………… .. 6. Cyphostemma
12. Tendrils simple or bifid; flowers functionally unisexual; stigma broadly 4- lobed on shon thick style or subsessile …… .. 4. Tetrastigma
+ Tendrils 1 – 2 x bifid; flowers bisexual; stigma minute on slender elongate style ……. 7. Cayratia (C. japonica)


 
Plant Taxonomy By O P. Sharma (1993)- Details


 
 
 
 
 

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